You are totally misrepresenting Nick Lane for the third time. He specifically rejects the "metabolism first" position and the alternative and lays out at great length why it is a "barren debate".
You claimed before that he said religion is of the greatest inventions of evolution - he did not.
Then you claimed that "the black hole at the heart of biology" is a metaphysical one about what makes us human - he says no such thing.
Now you have made it a hat trick by distorting the central premise of his ideas.