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Posts by hawkaw
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WT Child Abuse Coverup Reported Uzzah - 2nd Court Ruling in Kilbreath case
by hawkaw inrecall uzzah's original post on kilbreath v. sasketchewan (att.
gen)
then click on the url that states "kilbreath v. saskatchewan (attorney general)".
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WT Child Abuse Coverup Reported Uzzah - 2nd Court Ruling in Kilbreath case
by hawkaw inrecall uzzah's original post on kilbreath v. sasketchewan (att.
gen)
then click on the url that states "kilbreath v. saskatchewan (attorney general)".
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hawkaw
BTTT
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WT Child Abuse Coverup Reported Uzzah - 2nd Court Ruling in Kilbreath case
by hawkaw inrecall uzzah's original post on kilbreath v. sasketchewan (att.
gen)
then click on the url that states "kilbreath v. saskatchewan (attorney general)".
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hawkaw
Recall Uzzah's original post on Kilbreath v. Sasketchewan (Att. Gen)
http://www.jehovahs-witness.com/11/86969/1.ashx
Uzzah states
Essentially, a JW is accused of child abuse. The RCMP requests access to the local congregation files regarding complaints against the accused. Congregation refuses and sneds files to Bethel Legal. Legal Dept then claims Solicitor/Client privilege on the documents.
I believe the police (RCMP) won that case but had to wait for a deicsion on clerical privilege.
I did not realize but another decision was reported by the Court on April 5, 2005
Sadly the police lost and the search warrants were tossed since the police went apparently on a fishing expedition. They should ought to have had evidence about what was likely to be in the elders' papers but they never put it in their warrants. The Court never got to answer the privilege issue even though they did make note that a JP or Judge has to balance competing interests.
Anyway you can get to the case by going to the following URL here:
http://www.lawsociety.sk.ca/newlook/Library/fulltextnew.htm
In the Access Number line, Type in the Code: QB05122
Then use your mouse and click on "Submit Query". A new web page will show up.
Then click on the url that states "KILBREATH V. SASKATCHEWAN (ATTORNEY GENERAL)".
Below is the cases text. Hope it comes out okay in the post.
hawk
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KILBREATH V. SASKATCHEWAN (ATTORNEY GENERAL) QB05122 Date of Judgment: April 5, 2005 Number of Pages: 11 The judge feels that this judgment does not warrant publication. 2005 SKQB 165 Q.B.G. A.D. 2004
No. 526 J.C.B. IN THE COURT OF QUEEN'S BENCH JUDICIAL CENTRE OF BATTLEFORD
BETWEEN: PERRY KILBREATH , GORDON EMILSON, HAROLD McLAREN and LORNE VICKERS APPLICANTS - and - THE ATTORNEY GENERAL OF SASKATCHEWAN RESPONDENT David M. Gnam for the applicants K. Scott Bartlett for the respondent Micheal P. Hudec for the brother-in-law JUDGMENT APRIL 5, 2005 KRUEGER J. [1] On November 25, 2004, an application was made by Perry Kilbreath , Gordon Emilson, Harold McLaren and Lorne Vickers for an interim order sealing the documents seized (confidential record) pursuant to two search warrants, permitting examination of the officers who obtained the warrants and an order quashing the warrants or alternatively declaring the confidential record to be legally privileged. [2] By a fiat dated December 8, 2004, Wilkinson J., of this Court, granted leave to the applicants to examine the officers who obtained the warrants, granted leave to the respondent to cross-examine Perry Kilbreath on his affidavit, and ordered a hearing to determine whether the warrants ought to be quashed as invalid or as breaching the applicants' Charter rights of religious privilege. Much of the hearing before Madam Justice Wilkinson appears to have been dedicated to a discussion relating to solicitor/client privilege. That was no longer an issue at this hearing. No solicitor/client privilege exists or was claimed. BACKGROUND FACTS [3] Two search warrants were issued by a judge of the Provincial Court: 1. A search warrant issued on April 27, 2004, to conduct a search at Kingdom Hall, North Battleford, Saskatchewan, for a confidential record relating to sexual assaults on two females of the Jehovah's Witness faith by a male member of the Jehovah's Witness faith. The confidential record was thought to have been created during meetings between the accusing members, the accused member and elders of the faith. 2. A search warrant issued on October 1, 2004, to search for and seize the same confidential record then in the hands of solicitors in Toronto representing the applicant elders. That warrant was endorsed pursuant to s. 487(2) of the Criminal Code and seizure of the confidential record was effected in Ontario. [4] The search warrant that issued on April 24, 2004, was supported by the affidavit of Constable Wendell Theodore Houk, a member of the Royal Canadian Mounted Police, General Investigation Section of the Battlefords detachment. His affidavit is based on information obtained from the detachment records. On December 15, 2003, the two female Jehovah's Witness faith members (who are sisters) attended at the Battlefords detachment and complained that they had been separately sexually assaulted and harassed by their older sister's husband (brother-in-law) between the years 1981 and 1991. Without reference to specific times, both recalled occasions involving being touched or groped on the breast and in the vaginal area and having their brother-in-law rub his erect penis against them while hugging or otherwise contacting them. [5] The search warrant issued on October 1, 2004, was obtained on the basis of the affidavit of Constable Amber Lea Clark. She is also a member of the Royal Canadian Mounted Police, General Investigation Section at the Battlefords detachment. For the most part, her affidavit relies on the same information contained in the affidavit of Constable Houk. It updates some events. In particular, the affidavit of Constable Clark refers to telephone conversations that took place between Constable Houk and the applicant elder Perry Kilbreath advising him of the intent of the police to obtain a warrant and to seize the confidential record relating to the allegations made by the two sisters. During that telephone conversation Constable Houk attempted to arrange for a time and place during which the confidential record could be seized. The affidavit also refers to the mailing by the applicant, Emilson, of the confidential record to lawyers in Toronto. A telephone conversation is referred to between Constable Houk and the lawyer in Toronto relating to the possible sealing of the confidential record. [6] Shortly after September, 1991, the two sisters approached Jehovah's Witness elders in Lloydminster with their complaints. The meeting took place at the home of one of the sisters. Present at that meeting were the applicants Gordon Emilson and Harold McLaren, who are both elders of the Jehovah's Witness faith, Lloydminster congregation, the complainant sisters and their respective spouses. The accused brother-in-law was then resident in North Battleford and a member of the North Battleford Jehovah Witnesses' congregation. [7] Within a week of that meeting the two Lloydminster Jehovah's Witness elders Gordon Emilson and Harold McLaren met in North Battleford with elders Perry Kilbreath and Lorne Vickers and the accused brother-in-law, who was requested to be in attendance. During that meeting the accused brother-in-law acknowledged the complaints made against him by the two sisters and admitted to the acts complained of. He repented his wrongdoing. Because of his repentance it was resolved that the offending brother-in-law would not have his membership revoked. [8] The second meeting was described as the judicial action meeting. At that meeting the accused sinner, in this case the brother-in-law, was confronted in the absence of his accusers with their allegations. If denied, a face-to-face meeting could have resulted between the accusers and the accused. In this case acknowledgement and repentance made such a meeting unnecessary. At the conclusion of the second meeting all notes prepared by the four elders were destroyed. A record of the proceedings was prepared by elder Gordon Emilson and initialled or signed by the other elders. [9] The confidential record is an overview of the date, persons involved, accusations made (without details) and action taken. Some confidential records will also contain reference to counselling and scripture readings. In this case after the confidential record was signed by the elders, it was placed in an envelope, sealed and the names of the four elders were written on the outside of the envelope. That envelope was then locked in a safe in the Kingdom Hall at North Battleford. Only one of the four elders whose names appear on the outside of the envelope is entitled to open it. That envelope remained in the safe unaltered until removed by elder Gordon Emilson and sent to the lawyer in Toronto just prior to April 27, 2004. That is the confidential record that was seized pursuant to the search warrant issued on October 1, 2004. [10] About a week after the second meeting (judicial action meeting) a third meeting was held. At that meeting the accused brother-in-law and his spouse, the two sisters and their respective spouses, together with the four elders met for what was described as a reproof meeting. At that meeting no discussion relating to the allegations made or the response of the brother-in-law took place. The meeting consisted primarily of prayers and scripture readings. Its purpose was to provide some closure. The sisters were made aware that their complaints had been validated and the brother-in-law's repentance had prevented the revocation of his membership. [11] The sisters were not present when the confidential record was prepared by elder Gordon Emilson of the Jehovah's Witnesses' faith. They have not seen the confidential record and due to the strict confidentiality practiced by the elders are not aware of the contents of that record. [12] In his affidavit sworn April 27, 2004, Constable Houk attested to a belief that the confidential record will afford evidence of the offences complained of by the sisters. Likewise, in her affidavit sworn October 1, 2004, Constable Clark expressed the belief that the items being searched for (confidential record) will afford evidence of the offences alleged. The only possible source of information that could lead to such a belief is the sisters. Nowhere in the affidavits is the belief of either Constable Houk or Constable Clark attributed to the sisters or any other named source. The sisters do not possess any knowledge of the contents of the confidential record. It is possible that they assumed from the reproof meeting that the confidential record may include details of their accusations and a confession by their brother-in-law. THE LAW [13] The relevant portions of the Criminal Code relating to search warrants are: 487.(1) A justice who is satisfied by information on oath in Form 1 that there are reasonable grounds to believe that there is in a building, receptacle or place (a) ... (b) anything that there are reasonable grounds to believe will afford evidence with respect to the commission of an offence, or will reveal the whereabouts of a person who is believed to have committed an offence, against this Act or any other Act of Parliament. (c) ... (c.1) ... may at any time issue a warrant authorizing a peace officer or a public officer who has been appointed or designated to administer or enforce a federal or provincial law and whose duties include the enforcement of this Act or any other Act of Parliament and who is named in the warrant (d) to search the building, receptacle or place for any such thing and to seize it, and (e) subject to any other Act of Parliament, to, as soon as practicable, bring the thing seized before, or make a report in respect thereof to, the justice or some other justice for the same territorial division in accordance with section 489.1. (2) ... (3) ... (4) An endorsement that is made on a warrant as provided for in subsection (2) is sufficient authority to the peace officers or public officers to whom it was originally directed, and to all peace officers within the jurisdiction of the justice by whom it is endorsed, to execute the warrant and to deal with the things seized in accordance with section 489.1 or as otherwise provided by law. ANALYSIS
VALIDITY OF WARRANTS [14] In most cases where the search warrant is attacked that attack takes place during the trial of the accused. R. v. Tanner (1989), 46 C.C.C. (3d) 513 (Alta. C.A.); R. v. Williams (1987), 38 C.C.C. (3d) 319 (Y.T. C.A.); R. v. Jamieson (1989), 48 C.C.C. (3d) 287 (N.S .C.A.). Here, however, there is no accused and, accordingly, no possible trial. The search warrants were issued in aid of an ongoing police investigation. Neither the applicants nor their church are the subject of that investigation. There may never be charges laid. It is necessary to determine the validity of the warrants at this time in order that the investigation may go forward and the applicants have some finality. [15] As long as the Provincial Court judge acted judicially in concluding that there were reasonable grounds for issuing the warrants, the reviewing court should not substitute its view of the facts for that of the issuing judge. See: R. v. Jackson (1983), 9 C.C.C. (3d) 125 (B.C. C.A.). The issuing judge must exercise his/her judicial discretion based on reasonable and probable grounds that an offence has been committed or is suspected of having been committed and that the location of the search will provide evidence of the commission of an offence. R. v. Turcotte (1987), 39 C.C.C. (3d) 193 (Sask. C.A.). The issue in this case is whether the issuing Provincial Court judge had information that provided reasonable and probable grounds for the belief that the confidential record sought would provide evidence of the commission of an offence. [16] Each constable expressed a belief that the sought confidential record would afford such evidence. There is no basis for that belief. Both constables were aware from information received by them from the sisters that notes were prepared by the Jehovah's Witness elders at the meetings which the sisters attended. Nothing in either affidavit contains information as to what disclosures were made by the brother-in-law or what was recorded in the confidential record by the elders. [17] In R. v. Serendip Physiotherapy Clinic (2004), 189 C.C.C. (3d) 417 (Ont. C.A.), Rosenberg J.A. dealt with patient health records. At para. 35 he stated: It follows that the requirements of s. 487(1)(b), in particular the requirement that the officer provide information under oath of reasonable grounds to believe that the records sought "will" afford evidence with respect to the commission of an offence, strikes the proper balance even where the target of the search is the seizure of health records. By its terms, s. 487 precludes granting of a search warrant for the purposes of a fishing expedition or on the basis of mere suspicion. The same rationale, in my view, applies to religious counselling or spiritual records. Here the police had nothing more than a suspicion or speculation that the confidential record prepared by elder Gordon Emilson contained evidence with respect to the commission of an offence. Theirs was a fishing expedition. [18] The same conclusion was reached by the Saskatchewan Court of Appeal. In R. v. Pippin, [1994] S.J. No. 34 (Sask. C.A.), where Vancise J.A. stated at para. 12: The evidence before the justice of the peace is at best speculative. There is nothing in the third hand information which is sufficient to enable the justice of the peace, to test the reliability of the information, act judicially and issue the warrant. [19] Without considering whether the confidential record seized pursuant to the warrants contains confidential church information of a spiritual nature or whether there is a need of a balancing of the societal requirement to punish wrongdoers against the privilege of counselling procedures, the information in support of the warrants is wanting, to say the least. Cory J. in Canadian Broadcasting Corporation v. Lessard, [1991] 3 S.C.R. 421, at para. 14 stated: Certainly in every case the requirements of s. 487 of the Code must be met. However, this is not the end of the matter. Even after the statutory conditions have been met it may still be a difficult and complex process to determine whether a search warrant should be issued. Here there simply was insufficient information under oath of a credibly-based belief that the confidential record sought would provide evidence of an offence. The affidavits in support of the warrants do not justify their issuance. Accordingly, the warrants are invalid. RELIGIOUS PRIVILEGE OR CONFIDENTIALITY [20] Having found the search warrants to be invalid, the confidential records were unlawfully seized. They must be returned to the applicants either because the respondent has not indicated that anyone is likely to be charged or because it has not been shown that the confidential record contains evidence of an offence. Under the circumstances, it is not necessary to consider and balance the competing rights that seek protection of the public and breaches of the fundamental rights of those subject to the search. Nor is it necessary to examine the applicants' claim pursuant to the Charter of a valid religious privilege. [21] In passing, I would, however, comment that there is no merit in the argument that the affidavits failed to provide sufficient information to permit the Provincial Court judge to exercise a discretion by balancing the competing interests. There was sufficient information to identify the Jehovah's Witnesses as a religious congregation and the Kingdom Hall as a sacred place of worship. The confidential record sought was clearly the product of religious meetings. Even without using words such as "counselling", "spiritual", or "worship" the Provincial Court judge would have had no difficulty in recognizing the need to balance the competing interests in deciding whether to issue search warrants. [22] Counsel for the applicants also argued that there were alternative methods of obtaining the information gathered by the elders during their counselling meetings with the sisters and their brother-in-law. He referred, in particular, to the offer of counsel for the applicants to take the confidential record to the police in Ontario, have it sealed and made available for a voir dire to determine religious privilege. That was not a practical solution in the circumstances. There were and are no charges pending and no forum for a voir dire. [23] The two search warrants are quashed, the seized confidential record must be returned to the applicants in North Battleford. At the request of counsel on behalf of the brother-in-law, the hearing was held in camera. In keeping with that procedure, the public shall not have access to the court file or any documents which would tend to identify the sisters or their brother-in-law. -
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EARTH.GOOGLE - If you don't have it ... GET IT!
by hawkaw inthis may have already been posted but you have to see this website from google..
in some areas of the world you can zoom in to someones house on the earth..
amazing
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hawkaw
This may have already been posted but you have to see this website from GOOGLE..
In some areas of the world you can zoom in to someone’s HOUSE on the EARTH..
Amazing
To download go here
MUST HAVE HIGH SPEED CONNECTION FOR THE SOFTWARE TO FUNCTION...
Take care,
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Evolution of Bird, Mammals, Whales and Ants - with pictures
by hawkaw in4f), allin (1975, p. 3, fig.
procynosuchus modified from allin & hopson (1992, fig.
1); allin (1975, p. 3, fig.
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hawkaw
Thanks for the kind comment.
BTTT -
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Evolution of Bird, Mammals, Whales and Ants - with pictures
by hawkaw in4f), allin (1975, p. 3, fig.
procynosuchus modified from allin & hopson (1992, fig.
1); allin (1975, p. 3, fig.
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hawkaw
I saw this post on another site and thought I would post it here as it can help some (of course not all) understand the fossil record and how it relates to evolution. The poster, Ichneumon, really does a great job. As you will read he takes a few shots at creationists.
I still love looking at the feather remains in the below transitional vertebrate fossils.
hawk
Transitional Vertebrate Fossils FAQ
(Reptile to Bird etc.)...Theropod dinosaur to bird evolutionary transition:
The cladogram for the evolution of flight looks like this:(Note -- each name along the top is a known transitional fossil; and those aren't all that have been discovered.) Here's a more detailed look at the middle section:
Fossils discovered in the past ten years in China have answered most of the "which came first" questions about the evolution of birds from dinosaurs.
We now know that downy feathers came first, as seen in this fossil of Sinosauropteryx:
That's a close-up of downy plumage along the backbone. Here's a shot of an entire fossil
Sinosauropteryx was reptilian in every way, not counting the feathers. It had short forelimbs, and the feathers were all the same size. Presumably, the downy feathers evolved from scales driven by a need for bodily insulation.
Next came Protarchaeopteryx:
It had long arms, broad "hands", and long claws:
Apparently this species was driven by selection to develop more efficient limbs for grasping prey. One of the interesting things about this species is that the structure of the forelimb has been refined to be quite efficient at sweeping out quickly to grab prey, snap the hands together, then draw them back towards the body (mouth?). The specific structures in question are the semilunate carpal (a wrist bone), that moves with the hand in a broad, flat, 190 degree arc, heavy chest muscles, bones of the arm which link together with the wrist so as to force the grasping hands to spread out toward the prey during the forestroke and fold in on the prey during the upstroke. Not only is this a marvelously efficient prey-grabbing mechanism, but the same mechanism is at the root of the wing flight-stroke of modern birds. Evolution often ends up developing a structure to serve one need, then finds it suitable for adaptation to another. Here, a prey-grasping motion similar in concept to the strike of a praying mantis in a reptile becomes suitable for modifying into a flapping flight motion.
Additionally, the feathers on the hands and tail have elongated, becoming better suited for helping to sweep prey into the hands.
Next is Caudipteryx:
This species had hand and tail feathers even more developed than the previous species, and longer feathers, more like that of modern birds:
However, it is clear that this was still not a free-flying animal yet, because the forelimbs were too short and the feathers not long enough to support its weight, and the feathers were symmetrical (equal sized "fins" on each side of the central quill). It also had very reduced teeth compared to earlier specimens and a stubby beak:
But the elongation of the feathers indicates some aerodynamic purpose, presumably gliding after leaping (or falling) from trees which it had climbed with its clawed limbs, in the manner of a flying squirrel. Feathers which were developed "for" heat retention and then pressed into service to help scoop prey were now "found" to be useful for breaking falls or gliding to cover distance (or swooping down on prey?).
Next is Sinornithosaurus:
Similar to the preceding species, except that the pubis bone has now shifted to point to the back instead of the front, a key feature in modern birds (when compared to the forward-facing publis bone in reptiles). Here are some of the forearm feathers in detail:
Long feathers in detail:
Artists' reconstruction:
Next is Archaeopteryx:
The transition to flight is now well underway. Archaeopteryx has the reversed hallux (thumb) characteristic of modern birds, and fully developed feathers of the type used for flight (long, aligned with each other, and assymetrical indicating that the feathers have been refined to function aerodynamically). The feathers and limbs are easily long enough to support the weight of this species in flight. However, it lacks some structures which would make endurance flying more practical (such as a keeled sternum for efficient anchoring of the pectoral muscles which power the downstroke) and fused chest vertebrae. Archaeopteryx also retains a number of clearly reptilian features still, including a clawed "hand" emerging from the wings, small reptilian teeth, and a long bony tail. After the previous species' gliding abilities gave it an advantage, evolution would have strongly selected for more improvements in "flying" ability, pushing the species towards something more resembling sustained powered flight.
Next is Confuciusornis:
This species had a nearly modern flight apparatus. It also displays transitional traits between a reptilian grasping "hand" and a fully formed wing as in modern birds -- the outer two digits (the earlier species had three-fingered "hands") in Confuciusornis are still free, but the center digit has now formed flat, broad bones as seen in the wings of modern birds.
Additionally, the foot is now well on its way towards being a perching foot as in modern birds:
It also has a keeled sternum better suited for long flight, and a reduced number of vertebrae in the tail, on its way towards becoming the truncated tail of modern birds (which while prominent, is a small flap of muscle made to look large only because of the long feathers attached).
From this species it's only a small number of minor changes to finish the transition into the modern bird family.
(Hey, who said there are no transitional fossils? Oh, right, a lot of dishonest creationists. And there are a lot more than this, I've just posted some of the more significant milestones.)
There's been a very recent fossil find along this same lineage, too new for me to have found any online images to include in this article. And analysis is still underway to determine exactly where it fits into the above lineage. But it has well-formed feathers, which extend out from both the "arms" and the legs. Although it wasn't advanced enough to fully fly, the balanced feathering on the front and back would have made it ideally suited for gliding like a flying squirrel, and it may be another link between the stage where feathers had not yet been pressed into service as aerodynamic aids, and the time when they began to be used more and more to catch the air and developing towards a "forelimbs as wings" specialization.
So in short, to answer your question about how flight could have developed in birds, the progression is most likely some minor refinement on the following:
1. Scales modified into downy feathers for heat retention.
2. Downy feathers modified into "straight" feathers for better heat retention (modern birds still use their body "contour feathers" in this fashion).
3. Straight feathers modified into a "grasping basket" on the hands (with an accompanying increase in reach for the same purpose).4. Long limbs with long feathers refined to better survive falls to the ground.
5. "Parachute" feathers refined for better control, leading to gliding.
6. Gliding refined into better controlled, longer gliding.
7. Long gliding refined into short powered "hops".
8. Short powered flight refined into longer powered flight.
9. Longer powered flight refined into long-distance flying.Note that in each stage, the current configuration has already set the stage for natural selection to "prefer" individuals which better meet the requirements of the next stage. Evolution most often works like this; by taking some pre-existing ability or structure, and finding a better use for it or a better way to make it perform its current use.
archiopterix was or was not a fully developed Bird ...Explain these clearly reptilian features?
Premaxilla and maxilla are not horn-covered. This is posh talk for "does not have a bill."
[The above condensed from All About Archaeopteryx by Chris Nedin, which has far more information and quotes from primary research.Trunk region vertebra are free. In birds the trunk vertebrae are always fused.
Pubic shafts with a plate-like, and slightly angled transverse cross-section. A Character shared with dromaeosaurs but not with other dinosaurs or birds.
Cerebral hemispheres elongate, slender and cerebellum is situated behind the mid-brain and doesn't overlap it from behind or press down on it. This again is a reptilian feature. In birds the cerebral hemispheres are stout, cerebellum is so much enlarged that it spreads forwards over the mid-brain and compresses it downwards.
Neck attaches to skull from the rear as in dinosaurs not from below as in modern birds. The site of neck attachement (from below) is characteristic in birds, _Archaeopteryx_ does not have this character, but is the same as theropod dinosaurs.
Center of cervical vertebrae have simple concave articular facets. This is the same as the archosaur pattern. In birds the vertebrae are different, they have a saddle-shaped surface: "The most striking feature of the vertebrae is the simple disk-like facets of their centra, without any sign of the saddle-shaped articulations found in other birds" (de Beer 1954, p. 17).
Long bony tail with many free vertebrae up to tip (no pygostyle). Birds have a short tail and the caudal vertebrae are fused to give the pygostyle.
Premaxilla and maxilla bones bear teeth. No modern bird possess teeth.
Ribs slender, without joints or uncinate processes and do not articulate with the sternum. Birds have stout ribs with uncinate processes (braces between them) and articulate with the sternum.
Pelvic girdle and femur joint is archosaurian rather than avian (except for the backward pointing pubis as mentioned above).
The Sacrum (the vertebrae developed for the attachment of pelvic girdle) occupies 6 vertebra. This is the same as in reptiles and especially ornithipod dinosaurs. The bird sacrum covers between 11-23 vertebrae!
Metacarpals (hand) free (except 3rd metacarpal), wrist hand joint flexible. This is as in reptiles. In birds the metacarpals are fused together with the distal carpals in the carpo-metacarpus, wrist /hand fused.
Nasal opening far forward, separated from the eye by a large preorbital fenestra (hole). This is typical of reptiles, but not of birds.
Deltoid ridge of the humerus faces anteriorly as do the radial and ulnar condyles. Typical of reptiles but not found in birds.
Claws on 3 unfused digits. No modern adult bird has 3 claws, nor do they have unfused digits.
The fibula is equal in length to the tibia in the leg. This again is a typical character of reptiles. In birds the fibula is shortened and reduced. [When you eat a chicken drumstick, the fibula is the toothpick-like sliver of bone you find lying alongside the large "legbone", which is the tibia. Ich.]
Metatarsals (foot bones) free. In birds these are fused to form the tarsometatarsus.
Gastralia present. Gastralia are "ventral ribs," elements of dermal bone in the ventral wall of the abdomen. Typical of reptiles, they are absent in birds
If Evolution is True the Fossil record should have Millions of "Transitional Fossils"
It does.
Here are a few of the "non-existent" transitional fossils that some are incredibly ignorant about:
Reptile -> Mammal evolutionary transition:
The above is from 29+ Evidences for Macroevolution, which compiles several hundred transitional fossils, which is itself just a *SMALL* sampling of the ENORMOUS numbers of fine transitional sequences found in the fossil record and well known to anyone who has bothered to CRACK OPEN A BOOK -- or even do a websearch -- in the past 25 years or so... So what's the anti-evolutionists' excuse for remaining abysmally ignorant of such things, and repeatedly making the false claim that there are "no" transitional fossils, etc.?Example 2: reptile-mammals
Figure 1.4.1. The jaws of three vertebrates—mammal, therapsid, and pelycosaur. A side view of three idealized skulls of mammals, therapsids (mammal-like reptiles), and pelycosaurs (early reptiles). The figure shows the differences between mammal and reptilian jaws and ear-bone structures. The jaw joint is shown as a large black dot, the quadrate (mammalian anvil or incus) is in turquoise, the articular (mammalian hammer or malleus) is in yellow, and the angular (mammalian tympanic annulus) is in pink. Note how, in the reptile, the jaw joint is formed between the blue quadrate and the yellow articular (with the pink angular close by), and how, in the mammal, the jaw joint is formed between the squamosal above and the dentary below. In the reptile, the squamosal is just above and contacting the quadrate. Advanced therapsids have two jaw joints: a reptile-like joint and a mammal-like joint (Figure based on Kardong 2002, pp. 275, reproduced with permission from the publisher, Copyright © 2002 McGraw-Hill)
We also have an exquisitely complete series of fossils for the reptile-mammal intermediates, ranging from the pelycosauria, therapsida, cynodonta, up to primitive mammalia (Carroll 1988, pp. 392-396; Futuyma 1998, pp. 146-151; Gould 1990; Kardong 2002, pp. 255-275). As mentioned above, the standard phylogenetic tree indicates that mammals gradually evolved from a reptile-like ancestor, and that transitional species must have existed which were morphologically intermediate between reptiles and mammals—even though none are found living today. However, there are significant morphological differences between modern reptiles and modern mammals. Bones, of course, are what fossilize most readily, and that is where we look for transitional species from the past. Osteologically, two major striking differences exist between reptiles and mammals: (1) reptiles have at least four bones in the lower jaw (e.g. the dentary, articular, angular, surangular, and coronoid), while mammals have only one (the dentary), and (2) reptiles have only one middle ear bone (the stapes), while mammals have three (the hammer, anvil, and stapes) (see Figure 1.4.1).
Early in the 20 th century, developmental biologists discovered something that further complicates the picture. In the reptilian fetus, two developing bones from the head eventually form two bones in the reptilian lower jaw, the quadrate and the articular (see the Pelycosaur in Figure 1.4.1). Surprisingly, the corresponding developing bones in the mammalian fetus eventually form the anvil and hammer of the unique mammalian middle ear (also known more formally as the incus and malleus, respectively; see Figure 1.4.2) (Gilbert 1997, pp. 894-896). These facts strongly indicated that the hammer and anvil had evolved from these reptilian jawbones—that is, if common descent was in fact true. This result was so striking, and the required intermediates so outlandish, that many anatomists had extreme trouble imagining how transitional forms bridging these morphologies could have existed while retaining function. Young-earth creationist Duane Gish stated the problem this way:
"All mammals, living or fossil, have a single bone, the dentary, on each side of the lower jaw, and all mammals, living or fossil, have three auditory ossicles or ear bones, the malleus, incus and stapes. ... Every reptile, living or fossil, however, has at least four bones in the lower jaw and only one auditory ossicle, the stapes. ... There are no transitional fossil forms showing, for instance, three or two jawbones, or two ear bones. No one has explained yet, for that matter, how the transitional form would have managed to chew while his jaw was being unhinged and rearticulated, or how he would hear while dragging two of his jaw bones up into his ear." (Gish 1978, p. 80)
Figure 1.4.2. A comparison of the ears of reptiles and mammals. The reptile ear is shown on the left, the mammal ear on the right. As in Figure 1.4.1, the quadrate (mammalian anvil or incus) is in turquoise and the articular (mammalian hammer or malleus) is in yellow. The stapes is shown in brown. Note how the relative arrangement of these bones is similar in both taxa, in the order of inner ear-stapes-quadrate-articular.
Gish was incorrect in stating that there were no transitional fossil forms, and he has been corrected on this gaff numerous times since he wrote these words. However, Gish's statements nicely delineate the morphological conundrum at hand. Let's review the required evolutionary conclusion. During their evolution, two mammalian middle ear bones (the hammer and anvil, aka malleus and incus) were derived from two reptilian jawbones. Thus there was a major evolutionary transition in which several reptilian jawbones (the quadrate, articular, and angular) were extensively reduced and modified gradually to form the modern mammalian middle ear. At the same time, the dentary bone, a part of the reptilian jaw, was expanded to form the major mammalian lower jawbone. During the course of this change, the bones that form the hinge joint of the jaw changed identity. Importantly, the reptilian jaw joint is formed at the intersection of the quadrate and articular whereas the mammalian jaw joint is formed at the intersection of the squamosal and dentary (see Figure 1.4.1).
How could hearing and jaw articulation be preserved during this transition? As clearly shown from the many transitional fossils that have been found (see Figure 1.4.3), the bones that transfer sound in the reptilian and mammalian ear were in contact with each other throughout the evolution of this transition. In reptiles, the stapes contacts the quadrate, which in turn contacts the articular. In mammals, the stapes contacts the incus, which in turn contacts the malleus (see Figure 1.4.2). Since the quadrate evolved into the incus, and the articular evolved into the malleus, these three bones were in constant contact during this impressive evolutionary change. Furthermore, a functional jaw joint was maintained by redundancy—several of the intermediate fossils have both a reptilian jaw joint (from the quadrate and articular) and a mammalian jaw joint (from the dentary and squamosal). Several late cynodonts and Morganucodon clearly have a double-jointed jaw. In this way, the reptilian-style jaw joint was freed to evolve a new specialized function in the middle ear. It is worthy of note that some modern species of snakes have a double-jointed jaw involving different bones, so such a mechanical arrangement is certainly possible and functional.
Since Figure 1.4.3 was made, several important intermediate fossils have been discovered that fit between Morganucodon and the earliest mammals. These new discoveries include a complete skull of Hadrocodium wui (Luo et al. 2001) and cranial and jaw material from Repenomamus and Gobiconodon (Wang et al. 2001). These new fossil finds clarify exactly when and how the malleus, incus, and angular completely detached from the lower jaw and became solely auditory ear ossicles.
Recall that Gish stated: "There are no transitional fossil forms showing, for instance, three or two jawbones, or two ear bones" (Gish 1978, p. 80). Gish simply does not understand how gradual transitions happen (something he should understand, obviously, if he intends to criticize evolutionary theory). These fossil intermediates illustrate why Gish's statement is a gross mischaracterization of how a transitional form should look. In several of the known intermediates, the bones have overlapping functions, and one bone can be called both an ear bone and a jaw bone; these bones serve two functions. Thus, there is no reason to expect transitional forms with intermediate numbers of jaw bones or ear bones. For example, in Morganucodon, the quadrate (anvil) and the articular (hammer) serve as mammalian-style ear bones and reptilian jaw bones simultaneously. In fact, even in modern reptiles the quadrate and articular serve to transmit sound to the stapes and the inner ear (see Figure 1.4.2). The relevant transition, then, is a process where the ear bones, initially located in the lower jaw, become specialized in function by eventually detaching from the lower jaw and moving closer to the inner ear.
Figure 1.4.3. A comparison of the jawbones and ear-bones of several transitional forms in the evolution of mammals. Approximate stratigraphic ranges of the various taxa are indicated at the far left (more recent on top). The left column of jawbones shows the view of the left jawbone from the inside of the mouth. The right column is the view of the right jawbone from the right side (outside of the skull). As in Figure 1.4.1, the quadrate (mammalian anvil or incus) is in turquoise, the articular (mammalian hammer or malleus) is in yellow, and the angular (mammalian tympanic annulus) is in pink. For clarity, the teeth are not shown, and the squamosal upper jawbone is omitted (it replaces the quadrate in the mammalian jaw joint, and forms part of the jaw joint in advanced cynodonts and Morganucodon). Q = quadrate, Ar = articular, An = angular, I = incus (anvil), Ma = malleus (hammer), Ty = tympanic annulus, D = dentary. (Reproduced from Kardong 2002, pp. 274, with permission from the publisher, Copyright © 2002 McGraw-Hill)
Here's another look:
(The above is from The Fossil Record: Evolution or "Scientific Creation", which is yet ANOTHER source a lot of folks are not aware...)Mammal-Like Reptiles
As previously stated, a succession of transitional fossils exists that link reptiles (Class Reptilia) and mammals (Class Mammalia). These particular reptiles are classifie as Subclass Synapsida. Presently, this is the best example of th e transformation of one major higher taxon into another. The morphologic changes that took place are well documented by fossils, beginning with animals essentially 100% reptilian and resulting in animals essentially 100% mammalian. Therefore, I have chosen this as the example to summarize in more detail (Table 1, Fig. 1).
Skulls and jaws of synapsid reptiles and mammals; left column side view of skull; center column top view of skull; right column side view of lower jaw. Hylonomus modified from Carroll (1964, Figs. 2,6; 1968, Figs. 10-2, 10-5; note that Hylonomus is a protorothyrod, not a synapsid). Archaeothyris modified from Reisz (1972, Fig. 2). Haptodus modified from Currie (1977, Figs, 1a, 1b; 1979, Figs. 5a, 5b). Sphenacodo n modified from Romer & Price (1940, Fig. 4f), Allin (1975, p. 3, Fig. 16);note: Dimetrodon substituted for top view; modified from Romer & Price, 1940, pl. 10. Biarmosuchus modified from Ivakhnenko et al. (1997, pl. 65, Figs. 1a, 1B, 2); Alin & Hopson (1992; Fig. 28.4c); Sigogneau & Tchudinov (1972, Figs. 1, 15). Eoarctops modified from Broom (1932, Fig. 35a); Boonstra (1969, Fig. 18). Pristerognathus modified from Broom (1932, Figs 17a, b,c); Boonstra (1963, Fig. 5d). Procynosuchus modified from Allin & Hopson (1992, Fig. 28.4e); Hopson (1987, Fig. 5c); Brink (1963, Fig. 10a); Kemp (1979, Fig. 1); Allin (1975, p. 3, Fig. 14). Thrinaxodon modified from Allin & Hopson (1992, Fig. 28.4f);Parrington (1946, Fig. 1); Allin (1975, p. 3, Fig. 13). Probainognathus modified from Allin & Hopson (1992, Fig. 28.4g); Romer (1970, Fig. 1); Allin (1975, p. 3, Fig. 12). Morga nucodon modified from Kermack, Mussett, & Rigney (1981, Figs. 95, 99a; 1973, Fig. 7a); Allin (1975, p. 3, Fig. 11). Asioryctes modified from Carroll (1988, Fig. 20-3b). Abbreviations: ag = angular; ar = articular; cp = coronoid process; d = dentary; f = lateral temporal fenestra; j = jugal; mm = attachment site for mammalian jaw muscles; o = eye socket; po = post orbital; q = quadrate; rl = reflected lamina; sq = squamosal; ty = tympanic.
TAXONOMY LATERAL TEMPORAL FENESTRA LOWER JAW DENTARY TEETH LOWER JAW: POST- DENTARY BONES MIDDLE EAR & JAW ARTICULATION M: Early Placental mammals
Asioryctes
Upper CretaceousMerged with eye socket; cheek arch bowed out laterally 100% of jaw length is the den- tary; condylar process in contact with squamosal Fully differentiated teeth; incisors, canines, premolars; one tooth replacement No post-dentary bones 3 middle ear bones (stapes, incus, malleus) + tympanic; squamosal-dentary jaw joint L: "Pantothere" mammals
Amphitherium
Middle/Upper JurassicX 100% of jaw length is the den- tary; condylar process contacts squamosal Fully differentiated teeth; incisors, canines, premolars; one tooth replacement Post-dentary bones migrated to middle ear Probably 3 middle ear bones (stapes, incus, malleus) + tympanic; squamosal-dentary jaw joint K: Morganucodontid mammals
Morganucodon Upper Triassic & Lower JurassicMerged with eye socket; cheeck arch bowed out laterally 100% of jaw length is the den- tary; condylar process expanded posteriorly to make contact with squamosal Fully differentiated teeth; incisors, canines, premolars; one tooth replacement 20% of jaw length; reflected lamina decreased to narrow ribbon-like horseshoe Stapes extends from inner ear capsule to quadrate; quadrate tiny; both quadrate-articular and squamosal-dentary jaw joints J: Chiniquodontid cynodonts
Probainognathus
Middle TriassicMuch larger than eye socket; 40- 45% of skull length; expanded posterioirly, medially, & laterally; midline of skull narrow sagittal crest; chek arch bowed out laterally 95% of jaw length is the dentary; large coronoid process expanded posteriorly; condylar process expanded posteriorly Large single canine; cheek teeth multicusped; tooth replacement reduced 20% of jaw length; angular notch widened ventrally; width of main part of angular decreased; reflec - ted lamina decreased to narrow ribbon-like horseshoe Stapes extends from inner ear capsule to quadrate; quadrate tiny; quadrate-articular joint I:Galesaurid cynodonts
Thrinaxodon
Lower TriassicMuch larger than eye socket; 40% of skull length; expanded pos- terioirly, medially, & laterally; midline of skull narrow sagittal crest; chek arch bowed out laterally 85% of jaw length is the dentary; large coronoid process expanded to top of eye socket and pos- teriorly; jaw muscles attached to most of coronoid process Large single canine; cheek teeth multicusped; tooth replacement reduced 25% of jaw length; angular notch widened ventrally; width of reflec- ted lamina decreased; width of main part of angular decreased Stapes extends from inner ear capsule to quadrate; quadrate small; quadrate-articular jaw joint H: Procynosuchid cynodonts
Procynosuchus
upper Upper PermianMuch larger than eye socket; 40% of skull length; expanded pos- terioirly, medially, & laterally; midline of skull narrow sagittal crest; chek arch bowed out laterally 75-80% of jaw length is the den- tary; coronoid process expanded to near top of eye socket and posteriorly; jaw muscles attached to dorsal part of coronoid process Large single canine; cheek teeth multicusped 30% of jaw length; angular notch widened ventrally; width of reflected lamina decreased Stapes extends from inner ear capsule to quadrate; quadrate small; quadrate-articular jaw joint G: Early Therocephalians
Pristerognathus
lower Upper PermianLarger than eye socket; expanded posteriorly and medially; 30% of skull length 75-80% of jaw length is the den- tary; posterior end of dentary expanded posteriorly and dorsally into narrow blade-like coronoid process; rises to middle of eye socket Large single canine; other teeth simple cones. 35% of jaw length; angular notch deepened into a cleft; reflected lamina large, broad, blade-like Stapes extends from inner ear capsule to quadrate; quadrate small; quadrate-articular jaw joint F: Early Gorgonopsians
Eoarctops
lower Upper PermianSlightly larger than eye socket; expanded posteriorly and medially (minimal); 20-25% of skull length 65-75% of jaw length is the den- tary; posterior end of dentary slightly expanded posteriorly and dorsally as incipient coronoid process Large single canine; other teeth simple cones. 40% of jaw length; angular notch deepened into a cleft; reflected lamina large, broad, blade-like Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint E: Eotitanosuchians
Sphenacodon
Lower PermianSmall; slightly smaller than eye socket; slightly expanded posteriorly and medially 65-75% of jaw length is the den- tary; posterodorsal edge rises broadly but slightly above tooth row Large single canine; other teeth simple cones. 40% of jaw length; angular notch deepened into a cleft; reflected lamina large, broad, blade-like Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint D: Late sphenacodonts
Sphenacodon
Upper PennsylvanianSmall; smaller than eye socket; confined to one side of skull 65% of jaw length is the dentary; posterodorsal edge rises broadly but slightly above the tooth row Enlarged incipient canines; other teeth simple cones 60% of jaw length; venntral edge of angular notched ("angular notch") offsetting a short pro- tusion (reflected lamina) Stapes extends from inner ear capsule to quadrate; quadrate large and plate-like; quadrate- articular jaw joint C: Early spenacodonts
Haptodus
Upper PennsylvanianTiny; smaller than eye socket; confined to one side of skull 65-75% of jaw length is the den- tary; posterodorsal edge rises broadly but slightly above tooth row Undifferentiated; slightly enlarged incipient canines just behind nares 70% of jaw length; ventral edge of angular with shallow indentation Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint B: Early ophiacodonts
Archaothyris
upper Middle PennsylvanianTiny; smaller than eye socket; confined to one side of skull x Undifferentiated; slightly enlarged incipient canines just behind nares x Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint A: Protorothyrids
Hylonomus
lower Middle PennsylvanianAbsent 65-75% of jaw length is the den- tary; posterodorsal edge rises broadly but slightly above tooth row Undifferentiated; slightly enlarged incipient canines just behind nares 70% of jaw length; ventral edge of angular continuous Stapes extends from inner ear capsule to quadrate; quadrate- articular jaw joint Table 1: Morphology of synapsid reptiles and mammals (Note that Hylonomus is a protothyrid, not a synapsid). Data from references cited in text.
Modern reptiles and mammals are very distinctive, easily diagnosable, and do not intergrade. Reptiles are covered by scales, mammals by hair; reptiles are cold-blooded, mammals warm-blooded; reptiles do not suckle their young, mammals have mammary glands; reptiles have sprawling posture, mammals have upright posture. Most of these features are soft part anatomy or physiology that very rarely fossilize (although dinosaur skin impressions are known from Cretaceous sediments, and imprints of mammal hair are known from Eocene bats from Germany; Franzen, 1990). In the fossil record, we must look to skeletal features.
There are many skeletal features which allow us to distinguish the reptiles from the mammals (Carroll, 1988; Table 1, rows A, M). The single most important defining characteristic is the nature of the articulation of the lower jaw to the skull (Simpson, 1959). In reptiles, multiple bones comprise the lower jaw. A small bone at the posterior end of the lower jaw, the articular, articulates with the quadrate bone of the skull (Simpson, 1959; Carroll, 1988). In mammals, one large bone, the dentary, comprises the lower jaw. It articulates with the squamosal bone of the skull (Simpson, 1959; Carroll, 1988).
From comparative anatomy studies, it is certain that most of the bones of the reptiles and mammals are homologous (Crompton & Parker, 1978; Carroll, 1988). Of greatest importance, the middle ear bones of mammals (stapes, incus, malleus, and tympanic) are homologous with several of the skull and jaw bones of reptiles (stapes, quadrate, articular, and angular, respectively; Romer, 1956, p. 33-38, 1970a; Allin, 1975, 1986; Allin & Hopson, 1992; Crompton & Parker, 1978; Hopso n, 1987, 1994; Carroll, 1988). One group of reptiles, the synapsids (Subclass Synapsida), share with the mammals an additional homologous structure: the lateral temporal fenestra, which is an opening in the skull behind the eye socket at the triple junction between the squamosal, jugal , and post orbital bones (Broom, 1932; Frazetta, 1968; Kemp, 1982; Carroll, 1988). A band of bone composed of the jugal and the squamosal is adjacent to the lateral temporal fenestra (Broom, 1932; Kemp, 1982; Carroll, 1988). This is the cheek arch so characteristic of mammal skulls (Broom, 1932; Kemp, 1982; Carroll, 1988). Therefore, synapsids are commonly named the “mammal-like reptiles.”
The presence of diagnosable morphologic differences between reptiles (including the oldest reptiles and the oldest synapsids) and mammals distinguishes them as distinct taxa. This allows us to test evolution by looking for transitional forms between the two. Because many of the bones are homologous, we should find evidence illustrating how these bones were modified over time to become the new bones. Furthermore, these morphologic changes should happen in parallel and in geochronologic succession.
Synapsid reptiles inhabited Pangea from the Middle Pennsylvanian through the Early Jurassic (Kemp, 1982, 1985; Sloan, 1983; Carroll, 1988; Hopson, 1969, 1987, 1994; Hopson & Crompton, 1969; Hotton, et al., 1986; Crompton & Jenkins, 1973; Sidor & Hopson, 1998; Romer & Price, 1940; Broom, 1932; Boonstra, 1963, 1969, 1971; Tchudinov, 1983; Olson, 1944; Tatarinov, 1974; Vyushkov, 1955; Efremov, 1954). From the Early Permian through the Early Triassic, they were the largest and most abundant land animals (Sloan, 1983; Colbert, 1965). Though much less well known to the general public than dinosaurs, one of the “cereal box dinosaurs,” Dimetrodon (the sail-backed reptile), is a synapsid, not a dinosaur (Romer & Price, 1940; Carroll, 1988). The oldest mammals are Late Triassic (Kemp, 1982; Carroll, 1988). Below is a discussion of the geochronologic succession linking synapsids and mammals. The oldest reptiles (named protorothyrids; Carroll, 1964, 1988, p. 192-199) are from the lower Middle Pennsylvanian, and the oldest synapsids (Reisz, 1972) are from the upper Middle Pennsylvanian, both of Nova Scotia. Upper Pennsylvanian and Lower Permian forms are known primarily from the midcontinent and Permian Basin region of the United States (Romer & Price, 1940; Currie, 1977, 1979; Kemp, 1982; Sloan, 1983). The basal Upper Permian forms are known from Russia (Tchudinov, 1960, 1983; Efremov, 1954; Olson, 1962; Sigogneau & Tchudinov, 1972; Ivakhnenko et al., 1997). Most of the Upper Permian and Lower Triassic succession is known from southern Africa, especially the Great Karoo of South Africa (Broom, 1932; Boonstra, 1963, 1969, 1971; Hopson & Kitching, 1972; Kemp, 1982; Sloan, 1983). The Middle Triassic forms are from South America (Romer, 1969a, 1969b, 1970b, 1973; Romer & Lewis, 1973; Bonaparte & Barbarena, 1975), and the Upper Triassic and Lower Jurassic mammals are known from Eurasia (Kermack, Mussett, & Rigney, 1973, 1981; Kemp, 1982). Subsequent Mesozoic mammals are known from all over the world (Simpson, 1928; Lillegraven et al., 1979).
When placed in proper geochronologic succession, the synapsids naturally form a succession of taxa (genera and families) that progressively become more mammal-like and less reptile-like (Kemp, 1982, 1985; Sloan, 1983; Sidor & Hopson, 1998; Hopson, 1987, 1994). Morphologic changes, summarized in Table 1 and Figure 1, affect the entire skeletal anatomy of these animals, but are most clearly displayed in their skulls.
The lateral temporal fenestra increased in size from a tiny opening smaller than the eye socket to a giant opening occupying nearly half the length of the skull. Ultimately, it merged with the eye socket, thus producing the full development of the cheek arch so characteristic of mammals (Broom, 1932; Frazetta, 1968; Kemp, 1982; Sloan, 1983; Hopson, 1987, 1994; Carroll, 1988).
Successively, the relative proportion of the lower jaw comprised of the dentary bone (teeth-bearing bone) gradually increased until the entire lower jaw consisted of the dentary (Kemp, 1982; Sloan, 1983; Carroll, 1988; Hopson, 1987, 1994). In Pennsylvanian and Lower and basal Upper Permian synapsids, the postero-dorsal edge of the lower jaw rose broadly but only slightly above the level of the tooth row (Romer & Price, 1940; Currie, 1977, 1979; Ivakhnenko et al., 1997; Tchudinov, 1960, 1983; Efremov, 1954; Olson, 1962; Sigogneau & Tchudinov, 1972; Hopson, 1987, 1994). In succeeding forms, the posterior part of the dentary expanded dorsally and posteriorly as a blade-like process, and progressively became larger (Broom, 1932; Boonstra, 1963, 1969, 1971; Sigogneau, 1970; Brink, 1963; Kemp, 1979; Hopson, 1987, 1994), forming the coronoid process (Parrington, 1946; Fourie, 1974; Romer, 1969b, 1970b, 1973; Hopson, 1987, 1994) to which the mammalian-type jaw musculature is attached (Barghusen, 1968; Bramble, 1978; Crompton, 1972; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988). Concomitantly, the post-dentary bones progressively reduced in size (Allin, 1975; Crompton, 1972; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988; Hopson, 1987, 1994).
Beginning with the Upper Pennsylvanian sphenacodonts, a notch developed in the angular bone that offsets a projection, the reflected lamina (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Romer & Price, 1940; Currie, 1977, 1979; Kemp, 1982; Sloan, 1983; Carroll, 1988). The reflected lamina first became a large blade-like flange (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Ivakhnenko et al., 1997; Tchudinov, 1960, 1983; Efremov, 1954; Olson, 1962; Sigogneau & Tchudinov, 1972; Broom, 1932; Sigogneau, 1970; Boonstra, 1963, 1969, 1971), and then was progressively reduced to a delicate horseshoe-shaped bone (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Brink, 1963; Parrington, 1946; Fourie, 1974; Romer, 1969b, 1970b, 1973; Kermack, Mussett, & Rigney, 1973, 1981; Kemp, 1979, 1982; Sloan, 1983; Carroll, 1988).
Simultaneously, the quadrate progressively decreased in size (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Kemp, 1982; Sloan, 1983; Carroll, 1988). The articular did not decrease in size much, being small initially, but developed a downward-pointing prong (Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994; Kemp, 1982; Sloan, 1983; Carroll, 1988). In the synapsids, the lower jaw was hinged to the skull by the articular and quadrate bones (Crompton, 1972; Crompton & Parker, 1978; Allin, 1975; Allin & Hopson, 1992; Hopson, 1987, 1994). Thus they are classified as reptiles (Simpson, 1959; Kemp, 1982; Sloan, 1983; Carroll, 1988). As the quadrate and articular became smaller, they were relieved of their solid suture to the dentary and skull (Crompton, 1972; Allin, 1975, 1986; Allin & Hopson, 1992; Hopson, 1987, 1994; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988). A projection of the dentary extended posteriorly and made contact with the squamosal. Morganucodon possessed the mammalian dentary-squamosal jaw joint adjacent to the reptilian articular-quadrate jaw joint (Kermack, Mussett, & Rigney, 1973, 1981; Carroll, 1988). It is classified as the first mammal, but it is a perfect intermediate. Now that a new jaw joint was established, the quadrate and articular were subsequently relieved of that function (Crompton, 1972; Allin, 1975, 1986; Allin & Hopson, 1992; Hopson, 1987, 1994; Crompton & Parker, 1978; Kemp, 1982; Sloan, 1983; Carroll, 1988). Ultimately, in Middle and Upper Jurassic mammals, the tiny quadrate, articular, and ring-like angular migrated as a unit to the middle ear where they joined the stapes and became the incus, malleus, and tympanic bones (Allin, 197 5, 1986; Allin & Hopson, 1992; Hopson, 1987, 1994; Kemp, 1982; Sloan, 1983; Carroll, 1988).
Progressively, the teeth became differentiated. The large canines developed first, followed by the development of multicusped cheek teeth, reduced tooth replacement (Osborn & Crompton, 1973; Crompton & Parker, 1978), and finally full y differentiated incisors, canines, premolars, and molars with one tooth replacement during life (Kemp, 1982; Hopson, 1994).
Many other morphologic changes are documented in the fossil record. These demonstrate the morphologic and geochronologic succession from sprawling limb posture to upright limb posture of mammals (Jenkins, 1971; Romer & Lewis, 197 3; Kemp, 1982; Carroll, 1988; Hopson, 1994). As Jenkins (1971, p. 210) stated, “In details of morphology and function, the cynodont post-cranial skeleton should be regarded as neither ‘reptilian’ nor ‘mammalian’ but as transitional between the two classes .” Other changes have been adequately summarized elsewhere (Kemp, 1982; Sloan, 1983; Carroll, 1988; Hopson, 1994). Obviously, fundamental physiologic changes must have taken place as well, many of which are not directly preserved in the fossil record, though some can be inferred from the skeletal anatomy (Findlay, 1968; Kemp, 1982; Sloan, 1983, Carroll, 1988; Hopson, 1994).
This is well documented in the fossil record by a massive volume of incontrovertible data that cannot be explained away. Such large-scale, progressive, continuous, gradual, and geochronologically successive morphologic change (Sidor & Hopson, 1998) is descent with modification, and provides compelling evidence for evolution on a grand scale.
Fish to elephant evolutionary transition
Tell me, of any two consecutive fossils in the following list, do any differ so much from each other that anti-evolutionists wouldn't just write it off as "just adaptation", or "just microevolution"? [All of the listed specimens are actual fossils]
Fish to Amphibian transition:
(Most of the above text is from The Transitional Vertebrate Fossils FAQ, and is the result of hard work by Kathleen Hunt, who deserves the credit. I've just extracted the relevant individual portions and assembled them into one direct fish-to-elephant sequence.) If you like that, here are a few hundred more.1. Cheirolepis, (early Devonian, 400 million years ago) -- Primitive bony ray-finned fishes that gave rise to the vast majority of living fish. Heavy acanthodian-type scales, acanthodian-like skull, and big notocord.
2. Osteolepis (mid-Devonian, 390 million years ago) -- One of the earliest crossopterygian lobe-finned fishes, still sharing some characters with the lungfish (the other lobe-finned fishes). Had paired fins with a leg-like arrangement of major limb bones, capable of flexing at the "elbow", and had an early-amphibian-like skull and teeth.
3. Eusthenopteron, Sterropterygion (mid-late Devonian, 380 million years ago) -- Early rhipidistian lobe-finned fish roughly intermediate between early crossopterygian fish and the earliest amphibians. Skull very amphibian-like. Strong amphibian- like backbone. Fins very like early amphibian feet in the overall layout of the major bones, muscle attachments, and bone processes, with tetrapod-like tetrahedral humerus, and tetrapod-like elbow and knee joints. But there are no perceptible "toes", just a set of identical fin rays. Body & skull proportions rather fishlike.
4. Panderichthys, Elpistostege (mid-late Devonian, about 370 Mya) -- These "panderichthyids" are very tetrapod-like lobe-finned fish. Unlike Eusthenopteron, these fish actually look like tetrapods in overall proportions (flattened bodies, dorsally placed orbits, frontal bones! in the skull, straight tails, etc.) and have remarkably foot-like fins.
5. Obruchevichthys(middle Late Devonian, about 370 Mya -- Discovered in 1991 in Scotland, these are the earliest known tetrapod remains. The humerus is mostly tetrapod-like but retains some fish features. The discoverer, Ahlberg (1991), said: "It [the humerus] is more tetrapod-like than any fish humerus, but lacks the characteristic early tetrapod 'L-shape'...this seems to be a primitive, fish-like character....although the tibia clearly belongs to a leg, the humerus differs enough from the early tetrapod pattern to make it uncertain whether the appendage carried digits or a fin. At first sight the combination of two such extremities in the same animal seems highly unlikely on functional grounds. If, however, tetrapod limbs evolved for aquatic rather than terrestrial locomotion, as recently suggested, such a morphology might be perfectly workable."
6. Hynerpeton, Acanthostega, Ichthyostega (late Devonian, 360 Mya) -- A little later, the fin-to-foot transition was almost complete, and we have a set of early tetrapod fossils that clearly did have feet. The most complete are Ichthyostega, Acanthostega gunnari, and the newly described Hynerpeton bassetti (Daeschler et al., 1994). (There are also other genera known from more fragmentary fossils.) Hynerpeton is the earliest of these three genera (365 Ma), but is more advanced in some ways; the other two genera retained more fish- like characters longer than the Hynerpeton lineage did. Acanthostega still had internal gills, adding further support to the suggestion that unique tetrapod characters such as limbs with digits evolved first for use in water rather than for walking on land. Acanthostega also had a remarkably fish-like shoulder and forelimb. Ichthyostega was also very fishlike, retaining a fish-like finned tail, permanent lateral line system, and notochord. It turns out that Acanthostega's front foot had eight toes, and Ichthyostega's hind foot had seven toes, giving both feet the look of a short, stout flipper with many "toe rays" similar to fin rays. All you have to do to a lobe- fin to make it into a many-toed foot like this is curl it, wrapping the fin rays forward around the end of the limb. In fact, this is exactly how feet develop in larval amphibians, from a curled limb bud. Hynerpeton, in contrast, probably did not have internal gills and already had a well-developed shoulder girdle; it could elevate and retract its forelimb strongly, and it had strong muscles that attached the shoulder to the rest of the body (Daeschler et al., 1994).
7. Labyrinthodonts (eg Pholidogaster, Pteroplax) (late Dev./early Miss., 355 Mya) -- These larger amphibians still have some icthyostegid fish features, such as skull bone patterns, labyrinthine tooth dentine, presence & pattern of large palatal tusks, the fish skull hinge, pieces of gill structure between cheek & shoulder, and the vertebral structure. But they have lost several other fish features: the fin rays in the tail are gone, the vertebrae are stronger and interlocking, the nasal passage for air intake is well defined, etc.
Amphibian to Reptile transition:
8. Pholidogaster (Mississippian, about 330 Ma) -- A group of large labrinthodont amphibians, transitional between the early amphibians (the ichthyostegids, described above) and later amphibians such as rhachitomes and anthracosaurs.
9. Proterogyrinus (late Mississippian, 325 Mya) -- Classic labyrinthodont-amphibian skull and teeth, but with reptilian vertebrae, pelvis, humerus, and digits. Still has fish skull hinge. Amphibian ankle. 5-toed hand and a 2-3-4-5-3 (almost reptilian) phalangeal count.
10. Limnoscelis, Tseajaia (late Carboniferous, 300 Mya) -- Amphibians apparently derived from the early anthracosaurs, but with additional reptilian features: structure of braincase, reptilian jaw muscle, expanded neural arches.
11. Solenodonsaurus (mid-Pennsylvanian) -- An incomplete fossil, apparently between the anthracosaurs and the cotylosaurs. Loss of palatal fangs, loss of lateral line on head, etc. Still just a single sacral vertebra, though.
12. Hylonomus, Paleothyris (early Pennsylvanian) -- These are protorothyrids, very early cotylosaurs (primitive reptiles). They were quite little, lizard-sized animals with amphibian-like skulls (amphibian pineal opening, dermal bone, etc.), shoulder, pelvis, & limbs, and intermediate teeth and vertebrae. Rest of skeleton reptilian, with reptilian jaw muscle, no palatal fangs, and spool-shaped vertebral centra. Probably no eardrum yet.
13. Paleothyris (early Pennsylvanian) -- An early captorhinomorph reptile, with no temporal fenestrae at all.
14. Protoclepsydrops haplous (early Pennsylvanian) -- The earliest known synapsid reptile. Little temporal fenestra, with all surrounding bones intact. Had amphibian-type vertebrae with tiny neural processes. (reptiles had only just separated from the amphibians)
15. Clepsydrops (early Pennsylvanian) -- The second earliest known synapsid.
Reptile to Mammal transition:
16. Archaeothyris (early-mid Pennsylvanian) -- A slightly later ophiacodont. Small temporal fenestra, now with some reduced bones (supratemporal). Braincase still just loosely attached to skull. Slight hint of different tooth types. Still has some extremely primitive, amphibian/captorhinid features in the jaw, foot, and skull. Limbs, posture, etc. typically reptilian, though the ilium (major hip bone) was slightly enlarged.
17. Varanops (early Permian) -- Temporal fenestra further enlarged. Braincase floor shows first mammalian tendencies & first signs of stronger attachment to rest of skull (occiput more strongly attached). Lower jaw shows first changes in jaw musculature (slight coronoid eminence). Body narrower, deeper: vertebral column more strongly constructed. Ilium further enlarged, lower-limb musculature starts to change (prominent fourth trochanter on femur). This animal was more mobile and active. Too late to be a true ancestor, and must be a "cousin".
18. Haptodus (late Pennsylvanian) -- One of the first known sphenacodonts, showing the initiation of sphenacodont features while retaining many primitive features of the ophiacodonts. Occiput still more strongly attached to the braincase. Teeth become size-differentiated, with biggest teeth in canine region and fewer teeth overall. Stronger jaw muscles. Vertebrae parts & joints more mammalian. Neural spines on vertebrae longer. Hip strengthened by fusing to three sacral vertebrae instead of just two. Limbs very well developed.
19. Dimetrodon, Sphenacodon or a similar sphenacodont (late Pennsylvanian to early Permian, 270 Ma) -- More advanced pelycosaurs, clearly closely related to the first therapsids (next). Dimetrodon is almost definitely a "cousin" and not a direct ancestor, but as it is known from very complete fossils, it's a good model for sphenacodont anatomy. Medium-sized fenestra. Teeth further differentiated, with small incisors, two huge deep- rooted upper canines on each side, followed by smaller cheek teeth, all replaced continuously. Fully reptilian jaw hinge. Lower jaw bone made of multiple bones & with first signs of a bony prong later involved in the eardrum, but there was no eardrum yet, so these reptiles could only hear ground-borne vibrations (they did have a reptilian middle ear). Vertebrae had still longer neural spines (spectacularly so in Dimetrodon, which had a sail), and longer transverse spines for stronger locomotion muscles.
20. Biarmosuchia (late Permian) -- A therocephalian -- one of the earliest, most primitive therapsids. Several primitive, sphenacodontid features retained: jaw muscles inside the skull, platelike occiput, palatal teeth. New features: Temporal fenestra further enlarged, occupying virtually all of the cheek, with the supratemporal bone completely gone. Occipital plate slanted slightly backwards rather than forwards as in pelycosaurs, and attached still more strongly to the braincase. Upper jaw bone (maxillary) expanded to separate lacrymal from nasal bones, intermediate between early reptiles and later mammals. Still no secondary palate, but the vomer bones of the palate developed a backward extension below the palatine bones. This is the first step toward a secondary palate, and with exactly the same pattern seen in cynodonts. Canine teeth larger, dominating the dentition. Variable tooth replacement: some therocephalians (e.g Scylacosaurus) had just one canine, like mammals, and stopped replacing the canine after reaching adult size. Jaw hinge more mammalian in position and shape, jaw musculature stronger (especially the mammalian jaw muscle). The amphibian-like hinged upper jaw finally became immovable. Vertebrae still sphenacodontid-like. Radical alteration in the method of locomotion, with a much more mobile forelimb, more upright hindlimb, & more mammalian femur & pelvis. Primitive sphenacodontid humerus. The toes were approaching equal length, as in mammals, with #toe bones varying from reptilian to mammalian. The neck & tail vertebrae became distinctly different from trunk vertebrae. Probably had an eardrum in the lower jaw, by the jaw hinge.
21. Procynosuchus (latest Permian) -- The first known cynodont -- a famous group of very mammal-like therapsid reptiles, sometimes considered to be the first mammals. Probably arose from the therocephalians, judging from the distinctive secondary palate and numerous other skull characters. Enormous temporal fossae for very strong jaw muscles, formed by just one of the reptilian jaw muscles, which has now become the mammalian masseter. The large fossae is now bounded only by the thin zygomatic arch (cheekbone to you & me). Secondary palate now composed mainly of palatine bones (mammalian), rather than vomers and maxilla as in older forms; it's still only a partial bony palate (completed in life with soft tissue). Lower incisor teeth was reduced to four (per side), instead of the previous six (early mammals had three). Dentary now is 3/4 of lower jaw; the other bones are now a small complex near the jaw hinge. Jaw hinge still reptilian. Vertebral column starts to look mammalian: first two vertebrae modified for head movements, and lumbar vertebrae start to lose ribs, the first sign of functional division into thoracic and lumbar regions. Scapula beginning to change shape. Further enlargement of the ilium and reduction of the pubis in the hip. A diaphragm may have been present.
22. Dvinia [also "Permocynodon"] (latest Permian) -- Another early cynodont. First signs of teeth that are more than simple stabbing points -- cheek teeth develop a tiny cusp. The temporal fenestra increased still further. Various changes in the floor of the braincase; enlarged brain. The dentary bone was now the major bone of the lower jaw. The other jaw bones that had been present in early reptiles were reduced to a complex of smaller bones near the jaw hinge. Single occipital condyle splitting into two surfaces. The postcranial skeleton of Dvinia is virtually unknown and it is not therefore certain whether the typical features found at the next level had already evolved by this one. Metabolic rate was probably increased, at least approaching homeothermy.
23. Thrinaxodon (early Triassic) -- A more advanced "galesaurid" cynodont. Further development of several of the cynodont features seen already. Temporal fenestra still larger, larger jaw muscle attachments. Bony secondary palate almost complete. Functional division of teeth: incisors (four uppers and three lowers), canines, and then 7-9 cheek teeth with cusps for chewing. The cheek teeth were all alike, though (no premolars & molars), did not occlude together, were all single- rooted, and were replaced throughout life in alternate waves. Dentary still larger, with the little quadrate and articular bones were loosely attached. The stapes now touched the inner side of the quadrate. First sign of the mammalian jaw hinge, a ligamentous connection between the lower jaw and the squamosal bone of the skull. The occipital condyle is now two slightly separated surfaces, though not separated as far as the mammalian double condyles. Vertebral connections more mammalian, and lumbar ribs reduced. Scapula shows development of a new mammalian shoulder muscle. Ilium increased again, and all four legs fully upright, not sprawling. Tail short, as is necessary for agile quadrupedal locomotion. The whole locomotion was more agile. Number of toe bones is 2.3.4.4.3, intermediate between reptile number (2.3.4.5.4) and mammalian (2.3.3.3.3), and the "extra" toe bones were tiny. Nearly complete skeletons of these animals have been found curled up - a possible reaction to conserve heat, indicating possible endothermy? Adults and juveniles have been found together, possibly a sign of parental care. The specialization of the lumbar area (e.g. reduction of ribs) is indicative of the presence of a diaphragm, needed for higher O2 intake and homeothermy. NOTE on hearing: The eardrum had developed in the only place available for it -- the lower jaw, right near the jaw hinge, supported by a wide prong (reflected lamina) of the angular bone. These animals could now hear airborne sound, transmitted through the eardrum to two small lower jaw bones, the articular and the quadrate, which contacted the stapes in the skull, which contacted the cochlea. Rather a roundabout system and sensitive to low-frequency sound only, but better than no eardrum at all! Cynodonts developed quite loose quadrates and articulars that could vibrate freely for sound transmittal while still functioning as a jaw joint, strengthened by the mammalian jaw joint right next to it. All early mammals from the Lower Jurassic have this low-frequency ear and a double jaw joint. By the middle Jurassic, mammals lost the reptilian joint (though it still occurs briefly in embryos) and the two bones moved into the nearby middle ear, became smaller, and became much more sensitive to high-frequency sounds.
24. Cynognathus (early Triassic, 240 Ma; suspected to have existed even earlier) -- We're now at advanced cynodont level. Temporal fenestra larger. Teeth differentiating further; cheek teeth with cusps met in true occlusion for slicing up food, rate of replacement reduced, with mammalian-style tooth roots (though single roots). Dentary still larger, forming 90% of the muscle-bearing part of the lower jaw. TWO JAW JOINTS in place, mammalian and reptilian: A new bony jaw joint existed between the squamosal (skull) and the surangular bone (lower jaw), while the other jaw joint bones were reduced to a compound rod lying in a trough in the dentary, close to the middle ear. Ribs more mammalian. Scapula halfway to the mammalian condition. Limbs were held under body. There is possible evidence for fur in fossil pawprints.
25. Diademodon (early Triassic, 240 Ma; same strata as Cynognathus) -- Temporal fenestra larger still, for still stronger jaw muscles. True bony secondary palate formed exactly as in mammals, but didn't extend quite as far back. Turbinate bones possibly present in the nose (warm-blooded?). Dental changes continue: rate of tooth replacement had decreased, cheek teeth have better cusps & consistent wear facets (better occlusion). Lower jaw almost entirely dentary, with tiny articular at the hinge. Still a double jaw joint. Ribs shorten suddenly in lumbar region, probably improving diaphragm function & locomotion. Mammalian toe bones (2.3.3.3.3), with closely related species still showing variable numbers.
26. Probelesodon (mid-Triassic; South America) -- Fenestra very large, still separate from eyesocket (with postorbital bar). Secondary palate longer, but still not complete. Teeth double-rooted, as in mammals. Nares separated. Second jaw joint stronger. Lumbar ribs totally lost; thoracic ribs more mammalian, vertebral connections very mammalian. Hip & femur more mammalian.
27. Probainognathus (mid-Triassic, 239-235 Ma, Argentina) -- Larger brain with various skull changes: pineal foramen ("third eye") closes, fusion of some skull plates. Cheekbone slender, low down on the side of the eye socket. Postorbital bar still there. Additional cusps on cheek teeth. Still two jaw joints. Still had cervical ribs & lumbar ribs, but they were very short. Reptilian "costal plates" on thoracic ribs mostly lost. Mammalian #toe bones.
28. Pachygenelus, Diarthrognathus (earliest Jurassic, 209 Ma) -- These are trithelodontids. Inflation of nasal cavity, establishment of Eustachian tubes between ear and pharynx, loss of postorbital bar. Alternate replacement of mostly single- rooted teeth. This group also began to develop double tooth roots -- in Pachygenelus the single root of the cheek teeth begins to split in two at the base. Pachygenelus also has mammalian tooth enamel, and mammalian tooth occlusion. Double jaw joint, with the second joint now a dentary-squamosal (instead of surangular), fully mammalian. Incipient dentary condyle. Reptilian jaw joint still present but functioning almost entirely in hearing; postdentary bones further reduced to tiny rod of bones in jaw near middle ear; probably could hear high frequencies now. More mammalian neck vertebrae for a flexible neck. Hip more mammalian, with a very mammalian iliac blade & femur. Highly mobile, mammalian-style shoulder. Probably had coupled locomotion & breathing.
29. Sinoconodon (early Jurassic, 208 Ma) -- The next known very ancient proto-mammal. Eyesocket fully mammalian now (closed medial wall). Hindbrain expanded. Permanent cheekteeth, like mammals, but the other teeth were still replaced several times. Mammalian jaw joint stronger, with large dentary condyle fitting into a distinct fossa on the squamosal. This final refinement of the joint automatically makes this animal a true "mammal". Reptilian jaw joint still present, though tiny.
Proto-mammal to Placental Mammal transition:
30. Kuehneotherium (early Jurassic, about 205 Ma) -- A slightly later proto-mammal, sometimes considered the first known pantothere (primitive placental-type mammal). Teeth and skull like a placental mammal. The three major cusps on the upper & lower molars were rotated to form interlocking shearing triangles as in the more advanced placental mammals & marsupials. Still has a double jaw joint, though.
31. Eozostrodon, Morganucodon, Haldanodon (early Jurassic, ~205 Ma) -- A group of early proto-mammals called "morganucodonts". The restructuring of the secondary palate and the floor of the braincase had continued, and was now very mammalian. Truly mammalian teeth: the cheek teeth were finally differentiated into simple premolars and more complex molars, and teeth were replaced only once. Triangular- cusped molars. Reversal of the previous trend toward reduced incisors, with lower incisors increasing to four. Tiny remnant of the reptilian jaw joint. Once thought to be ancestral to monotremes only, but now thought to be ancestral to all three groups of modern mammals -- monotremes, marsupials, and placentals.
32. Peramus (late Jurassic, about 155 Ma) -- A "eupantothere" (more advanced placental-type mammal). The closest known relative of the placentals & marsupials. Triconodont molar has with more defined cusps. This fossil is known only from teeth, but judging from closely related eupantotheres (e.g. Amphitherium) it had finally lost the reptilian jaw joint, attaing a fully mammalian three-boned middle ear with excellent high-frequency hearing. Has only 8 cheek teeth, less than other eupantotheres and close to the 7 of the first placental mammals. Also has a large talonid on its "tribosphenic" molars, almost as large as that of the first placentals -- the first development of grinding capability.
33. Endotherium (very latest Jurassic, 147 Ma) -- An advanced eupantothere. Fully tribosphenic molars with a well- developed talonid. Known only from one specimen. From Asia; recent fossil finds in Asia suggest that the tribosphenic molar evolved there.
34. Vincelestes neuquenianus (early Cretaceous, 135 Ma) -- A probably-placental mammal with some marsupial traits, known from some nice skulls. Placental-type braincase and coiled cochlea. Its intracranial arteries & veins ran in a composite monotreme/placental pattern derived from homologous extracranial vessels in the cynodonts. (Rougier et al., 1992)
35. Kennalestes and Asioryctes (late Cretaceous, Mongolia) -- Small, slender animals; eyesocket open behind; simple ring to support eardrum; primitive placental-type brain with large olfactory bulbs; basic primitive tribosphenic tooth pattern. Canine now double rooted. Still just a trace of a non-dentary bone, the coronoid, on the otherwise all-dentary jaw. "Could have given rise to nearly all subsequent placentals." says Carroll (1988).
Placental mammal to elephant transition:
36. Protungulatum (latest Cretaceous) -- Transitional between earliest placental mammals and the condylarths (primitive, small hoofed animals). These early, simple insectivore- like small mammals had one new development: their cheek teeth had grinding surfaces instead of simple, pointed cusps. They were the first mammal herbivores. All their other features are generalized and primitive -- simple plantigrade five-toed clawed feet, all teeth present (3:1:4:3) with no gaps, all limb bones present and unfused, pointy-faced, narrow small brain, eyesocket not closed.
37. Minchenella or a similar condylarth (late Paleocene) -- Known only from lower jaws. Has a distinctive broadened shelf on the third molar.
38. Phenacolophus (late Paleocene or early Eocene) -- An early embrithopod (very early, slightly elephant-like condylarths), thought to be the stem-group of all elephants.
39. Pilgrimella (early Eocene) -- An anthracobunid (early proto-elephant condylarth), with massive molar cusps aligned in two transverse ridges.
40. Unnamed species of proto-elephant (early Eocene) -- Discovered recently in Algeria. Had slightly enlarged upper incisors (the beginnings of tusks), and various tooth reductions. Still had "normal" molars instead of the strange multi-layered molars of modern elephants. Had the high forehead and pneumatized skull bones of later elephants, and was clearly a heavy-boned, slow animal. Only one meter tall.
41. Moeritherium, Numidotherium, Barytherium (early-mid Eocene) -- A group of three similar very early elephants. It is unclear which of the three came first. Pig-sized with stout legs, broad spreading feet and flat hooves. Elephantish face with the eye set far forward & a very deep jaw. Second incisors enlarged into short tusks, in upper and lower jaws; little first incisors still present; loss of some teeth. No trunk.
42. Paleomastodon, Phiomia (early Oligocene) -- The first "mastodonts", a medium-sized animals with a trunk, long lower jaws, and short upper and lower tusks. Lost first incisors and canines. Molars still have heavy rounded cusps, with enamel bands becoming irregular. Phiomia was up to eight feet tall.
43. Gomphotherium (early Miocene) -- Basically a large edition of Phiomia, with tooth enamel bands becoming very irregular. Two long rows cusps on teeth became cross- crests when worn down. Gave rise to several families of elephant- relatives that spread all over the world. From here on the elephant lineages are known to the species level.
44a. The mastodon lineage split off here, becoming more adapted to a forest browser niche, and going through Miomastodon (Miocene) and Pliomastodon (Pliocene), to Mastodon (or "Mammut", Pleistocene).
44b. Meanwhile, the elephant lineage became still larger, adapting to a savannah/steppe grazer niche:
45. Stegotetrabelodon (late Miocene) -- One of the first of the "true" elephants, but still had two long rows of cross-crests, functional premolars, and lower tusks. Other early Miocene genera show compression of the molar cusps into plates (a modern feature ), with exactly as many plates as there were cusps. Molars start erupting from front to back, actually moving forward in the jaw throughout life.
46. Primelephas (latest Miocene) -- Short lower jaw makes it look like an elephant now. Reduction & loss of premolars. Very numerous plates on the molars, now; we're now at the modern elephants' bizarre system of one enormous multi-layered molar being functional at a time, moving forward in the jaw.
47. Primelephas gomphotheroides (mid-Pliocene) -- A later species that split into three lineages, Loxodonta, Elephas, and Mammuthus:
- Loxodonta adaurora (5 Ma). Gave rise to the modern African elephant Loxodonta africana about 3.5 Ma.
- Elephas ekorensis (5 Ma), an early Asian elephant with rather primitive molars, clearly derived directly from P. gomphotheroides. Led directly to:
- Elephas recki, which sent off one side branch, E. hydrusicus, at 3.8 Ma, and then continued changing on its own until it became E. iolensis.
- Elephas maximus, the modern Asian elephant, clearly derived from
- E. hysudricus. Strikingly similar to young E. hysudricus animals. Possibly a case of neoteny (in which "new" traits are simply juvenile features retained into adulthood).
- Mammuthus meridionalis, clearly derived from P. gomphotheroides. Spread around the northern hemisphere. In Europe, led to M. armeniacus/trogontherii, and then to M. primigenius. In North America, led to M. imperator and then M. columbi.
Species-species transitions among the elephants:
- Maglio (1973) studied Pleistocene elephants closely. Overall, Maglio showed that at least 7 of the 17 Quaternary elephant species arose through smooth anagenesis transitions from their ancestors. For example, he said that Elephas recki "can be traced through a progressive series of stages...These stages pass almost imperceptibly into each other....In the late Pleistocene a more progressive elephant appears which I retain as a distinct species, E. iolensis, only as a matter of convenience. Although as a group, material referred to E. iolensis is distinct from that of E. recki, some intermediate specimens are known, and E. iolensis seems to represent a very progressive, terminal stage in the E. recki specific lineage."
- Maglio also documented very smooth transitions between three Eurasian mammoth species: Mammuthus meridionalis --> M. armeniacus (or M. trogontherii) --> M. primigenius.
- Lister (1993) reanalyzed mammoth teeth and confirmed Maglio's scheme of gradual evolution in European mammoths, and found evidence for gradual transitions in the North American mammoths too.
Similar fossil sequences can be listed for the majority of other major-group transitions.
(Did I hear a creationist in the back row say something about "no transitional fossils?")
Note that the changes between any two sequential transitionals are small enough that most creationists would write them off as only "microevolution" -- and yet those 50-or-so "microevolutionary" steps turn a fish into an elephant, which even the most stubborn creationist would have to concede is "macroevolution".
Once you've answered the first question, here's a second one for you: If evolutionary common descent *hasn't* actually happened -- if the different animal "kinds" were just *poofed* into existence fully-formed -- then why is it possible to order known fossils into such a smooth "transitional" chain *at all*, in a way that makes sense and is chronologically, morphologically, genetically, paleontologically, geographically (etc. etc.) consistent with the (allegedly) "non-existent" evolutionary transitions? And no, it's not possible to assemble a sequence of fossils in just any damned order you want, so don't try *that* excuse -- even evolutionary biologists aren't capable of putting together a transitional fossil sequence "showing", say, a cat evolving into a bird, or a butterfly into a bat. Please explain.
Evolution of whales from terrestrial mammals
(From Plagiarized Errors and Molecular Genetics)
A particularly impressive example of shared retroposons has recently been reported linking cetaceans (whales, dolphins and porpoises) to ruminants and hippopotamuses, and it is instructive to consider this example in some detail. Cetaceans are sea-living animals that bear important similarities to land-living mammals; in particular, the females have mammary glands and nurse their young. Scientists studying mammalian anatomy and physiology have demonstrated greatest similarities between cetaceans and the mammalian group known as artiodactyls (even-toed ungulates) including cows, sheep, camels and pigs. These observations have led to the evolutionist view that whales evolved from a four-legged artiodactyl ancestor that lived on land. Creationists have capitalized on the obvious differences between the familiar artiodactyls and whales, and have ridiculed the idea that whales could have had four-legged land-living ancestors. Creationists who claim that cetaceans did not arise from four-legged land mammals must ignore or somehow dismiss the fossil evidence of apparent whale ancestors looking exactly like one would predict for transitional species between land mammals and whales--with diminutive legs and with ear structures intermediate between those of modern artiodactyls and cetaceans (Nature 368:844,1994; Science 263: 210, 1994). (A discussion of fossil ancestral whale species with references may be found at http://www.talkorigins.org/faqs/faq-transitional/part2b.html#ceta) Creationists must also ignore or dismiss the evidence showing the great similarity between cetacean and artiodactyl gene sequences (Molecular Biology & Evolution 11:357, 1994; ibid 13: 954, 1996; Gatesy et al, Systematic Biology 48:6, 1999).
Recently retroposon evidence has solidified the evolutionary relationship between whales and artiodactyls. Shimamura et al. (Nature 388:666, 1997; Mol Biol Evol 16: 1046, 1999; see also Lum et al., Mol Biol Evol 17:1417, 2000; Nikaido and Okada, Mamm Genome 11:1123, 2000) studied SINE sequences that are highly reduplicated in the DNA of all cetacean species examined. These SINES were also found to be present in the DNA of ruminants (including cows and sheep) but not in DNA of camels and pigs or more distantly related mammals such as horse, elephant, cat, human or kangaroo. These SINES apparently originated in a specific branch of ancestral artiodactyls after this branch diverged from camels, pigs and other mammals, but before the divergence of the lines leading to modern cetaceans, hippopotamus and ruminants. (See Figure 5.) In support of this scenario, Shimamura et al. identified two specific insertions of these SINES in whale DNA (insertions B and C in Figure 5) and showed that in DNA of hippopotamus, cow and sheep these same two sites contained the SINES; but in camel and pig DNA the same sites were "empty" of insertions. More recently, hippopotamus has been identified as the closest living terrestrial relative of cetaceans since hippos and whales share retroposon insertions (illustrated by D and E in Figure 5) that are not found in any other artiodactyls (Nikaido et al, PNAS 96:10261, 1999). The close hippo-whale relationship is consistent with previously reported sequence similarity comparisons (Gatesy, Mol Biol Evol 14:537, 1997) and with recent fossil finds (Gingerich et al., Science 293:2239, 2001; Thewissen et al., Nature 413:277, 2001) that resolve earlier paleontological conflicts with the close whale-hippo relationship. (Some readers have wondered: if ruminants are more closely related to whales than to pigs and camels, why are ruminants anatomically more similar to pigs and camels than they are to whales? Apparently this results from the fact that ruminants, pigs and camels changed relatively little since their last common ancestor, while the cetacean lineage changed dramatically in adapting to an aquatic lifestyle, thereby obliterating many of the features -- like hooves, fur and hind legs -- that are shared between its close ruminant relatives and the more distantly related pigs and camels. This scenario illustrates the fact that the rapid evolutionary development of adaptations to a new niche can occur through key functional mutations, leaving the bulk of the DNA relatively unchanged. The particularly close relationship between whales and hippos is consistent with several shared adaptations to aquatic life, including use of underwater vocalizations for communication and the absence of hair and sebaceous glands.) Thus, retroposon evidence strongly supports the derivation of whales from a common ancestor of hippopotamus and ruminants, consistent with the evolutionary interpretation of fossils and overall DNA sequence similarities. Indeed, the logic of the evidence from shared SINEs is so powerful that SINEs may be the best available characters for deducing species relatedness (Shedlock and Okada, Bioessays 22:148, 2000), even if they are not perfect (Myamoto, Curr. Biology 9:R816, 1999).Figure 5. Specific SINE insertions can act as "tracers" that illuminate phylogenetic relationships. This figure summarizes some of the data on SINEs found in living artiodactyls and shows how the shared insertions can be interpreted in relation to evolutionary branching. A specific SINE insertion event ("A" in the Figure) apparently occurred in a primitive common ancestor of pigs, ruminants, hippopotamus and cetaceans, since this insertion is present in these modern descendants of that common ancestor; but it is absent in camels, which split off from the other species before this SINE inserted. More recent insertions B and C are present only in ruminants, hippopotamus and cetaceans. Insertions D and E are shared only by hippopotamus and cetaceans, thereby identifying hippopotamus as the closest living relative of cetaceans (at least among the species examined in these studies). SINE insertions F and G occurred in the ruminant lineage after it diverged from the other species; and insertions H and I occurred after divergence of the cetacean lineage. That's just a quick layman-level overview of *one* of the many ways that whale evolution has been verified. For more technical examinations along several independent lines of evidence, see for example:
SINE Evolution, Missing Data, and the Origin of WhalesPhylogenetic relationships among cetartiodactyls based on insertions of short and long interpersed elements: Hippopotamuses are the closest extant relatives of whalesEvidence from Milk Casein Genes that Cetaceans are Close Relatives of Hippopotamid ArtiodactylsAnalyses of mitochondrial genomes strongly support a hippopotamus±whale cladeA new, diminutive Eocene whale from Kachchh (Gujarat, India) and its implications for locomotor evolution of cetaceansA new Eocene archaeocete (Mammalia, Cetacea) from India and the time of origin of whalesMysticete (Baleen Whale) Relationships Based upon the Sequence of the Common Cetacean DNA Satellite1The Mitochondrial Genome of the Sperm Whale and a New Molecular Reference for Estimating Eutherian Divergence Dates Limbs in whales and limblessness in other vertebrates: mechanisms of evolutionary and developmental transformation and loss Eocene evolution of whale hearingNovel Phylogeny of Whales Revisited but Not RevisedLand-to-sea transition in early whales: evolution of Eocene Archaeoceti (Cetacea) in relation to skeletal proportions and locomotion of living semiaquatic mammalsSubordinal artiodactyl relationships in the light of phylogenetic analysis of 12 mitochondrial protein-coding genesNew Morphological Evidence for the Phylogeny of Artiodactyla, Cetacea, and MesonychidaeCetacean SystematicsLIKELIHOOD ESTIMATION OF THE TIME OF ORIGIN OF CETACEA AND THE TIME OF DIVERGENCE OF CETACEA AND ARTIODACTYLAPhylogenetic Relationships of Artiodactyls and Cetaceans as Deduced from the Comparison of Cytochrome b and 12s rRNA Mitochondrial SequencesMolecular evolution of mammalian ribonucleases
And much, much more.
Evolution of army ants
I got curious about the actual study, so after reading the Cornell press release, I went and bought a copy of the actual study from PNAS Online ("Evolution of the army ant syndrome: The origin and long-term evolutionary stasis of a complex of behavioral and reproductive adaptations").
The first thing I found is that there's a reason why creationists really ought to break their habit of learning their "science" from a) creationist sites or b) press releases (and why evolutionists more often than not read the primary literature, which is something I've almost never seen a creationist doing).
The press release did indeed say that "because since the reign of the dinosaurs, about 100 million years ago, army ants in essence have not changed a bit". But this is just the reporter's piss-poor misunderstanding (and therefore misrepresentation) of what the study *actually* determined. Nowhere in the study itself is there any hint of a claim that army ants "haven't changed a bit" in the past 100 million years.
What it *does* say is:
The army ant syndrome of behavioral and reproductive traits (obligate collective foraging, nomadism, and highly specialized queens) has allowed these organisms to become the premiere social hunters of the tropics [...] Results strongly indicate that the suite of behavioral and reproductive adaptations found in army ants throughout the world is inherited from a unique common ancestor [...] Because no known army ant species lacks any component of the army ant syndrome, this group represents an extraordinary case of long-term evolutionary stasis in these adaptations.
In other words, the three *characteristics* which make an army ant what it is (1. foraging in groups -- most non-army ants use scouts, 2. nomadic lifestyle -- most non-army ants nest in one place, 3. flightless queens which can pump out *millions* of eggs) came into being 100 million years ago and have persisted ever since. Needless to say, that's quite a different thing than the much more general "army ants haven't changed a bit" claim the reporter made.
Nor is the press release's headline accurate or supported by the study ("Army ants, as voracious as ever, have defied evolution for 100 million years, Cornell entomologist finds ").
There's nothing in the study about "defying evolution". The only thing I could find which the reporter might have (very badly) mistaken for such a claim is where the author points out that previous *assumptions* (which were recognized to be no more than assumptions) were that "old world" and "new world" army ants may have evolved separately from non-army ancestors on their respective continents. Instead, this study finds, there was a single army-ant ancestor and all modern army ant families/species descended from them. So the results of this study "defy" previous presumptions about how army ants may have evolved, but hardly "defy" evolution itself.
Someone shoot that reporter...
Meanwhile, the study's findings are interesting in their own right, and add yet more data to the massive mountains of hard evidence *for* evolution (which insulting cartoons by our resident creationists do nothing to refute).
Using a variety of measures (DNA base-pair sequences consisting of three nuclear and one mitochondrial gene totalling 3538 basepairs from each of the 49 extant ant species, fossil evidence, and 116 morphological metrics), the author's mathematical analysis produced a cladistic tree for both army ants and many non-army ant species as follows:
The letters (A-H) indicate points in time where the subsequent "branches" are known to have already existed, because representatives from each "branch" have been disocovered in the fossil record.
The branches marked with "*" are branches where the ML tree analysis produced results with "a posterior probability of >95% after independent Bayesian phylogenetic analysis".
As described in the press release, this does indeed clearly indicate that all modern army ants (species shown in bold type) descended from a common ancestor, instead of from two or more common ancestors which were themselves not army ants.
It's also interesting to note that all the "old world" (OW) and "new world" (NW) army ants are separate branches of the oldest split of the army ant family tree. This demonstrates that, as previously presumed, the lifestyle of the army ant (especially, wingless queens) precludes any cross-continental "crossovers", where some species had (during the last 100 million years) managed to travel from one continent to another and take up new residence there.
This correct presumption -- along with the incorrect presumption that army ants had appeared more recently than 100 million years ago -- was the basis for the original assumption that old-world and new-world army ants had perhaps evolved independently (on their respective continents).
Instead, this DNA and morphological analysis (which, by the way, does *not* depend on a "genetic clock") strongly indicates that army ants first arose approximately 105 million years ago.
The reason that this is such an interesting result is that it *very* closely matches the known time of existence of the Cretaceous super-continent of Gondwana, *and* the time of the old-world/new-world army ant split matches the known time of the break-up of that supercontinent into separate continents which contain what is now South America (on one side) and Africa (on the other), the respective homes of the new-world and old-world army ants.
In other words, the analysis strongly matches an evolutionary model in many different ways, including several I haven't even mentioned here.
First, the fact that such a "family tree" works out *at all* is strong evidence that evolution has actually taken place. If instead ants of various species and/or "kinds" had been separately created, there's no reason at all that their DNA details *and* their fossil traces *and* their morphological details would so neatly fit a timewise evolutionary tree of common descent *at all*. For just one example, if the species at the top of the tree and the bottom of the tree shared a characteristic gene sequence, while the other species didn't (because, say, God felt they each would benefit from it), then the entire tree structure would be blatantly violated. Instead, every time DNA/morphological data is analyzed in this way, even across widely divergent species like cows and giraffes and whales, an implied "tree of common descent" is inarguably implied by the evidence.
Second, in this case, the "family tree" implied by the evidence "just happens" to *exactly* match geologic events which would be expected to explain parts of the tree if it came about via evolution. For example, if all modern army ants had a common ancestor, then at some point in time the ancestral army ant must have arisen at a particular geographic location (obviously). This would be a problem if, for example, the data implied that this happened at a time before ants existed at all, or after army ants were known to exist in fossils, etc. And yet, when the available evidence is objectively analyzed by a mathematical algorithm with no ideological ax to grind, the results beautifully match an evolutionary origin consistent with the known fossil record, timewise.
Furthermore, red flags would be raised if the time-and-place of the calculated origin happened to fall in a place where army ants would be highly unlikely to have gotten from their point of origin to the separate continents where they are seen today (e.g. South America and Africa). But lo and behold, the analysis shows the time-and-place of the calculated origin to be at a time when those two continents were known to be joined.
Furthermore, the calculated split between old-world and new-world army ants is found to fall at a time when the continents themselves split apart, perfectly explaining how and why the populations on each new continent, now isolated, should (and thus did) diverge into families of species which evolved in unrelated directions from each other (thus forming species that, while all still army ants in lifestyle, show characteristic differences).
And so on and so on.
Again and again, every time studies and analyses like this -- and every other conceivable type -- are performed, the results "just happen" to fall in a way that makes perfect sense if modern (and fossil) life had arisen from earlier life forms in a common-descent, evolutionary process, like individual jigsaw puzzle pieces all of which form a smooth, coherent picture (albeit with some pieces still not yet discovered) where all the pieces found so far all mesh smoothly with their neighbors.
If evolution is *not* true, why does the jigsaw puzzle formed by the mountains of evidence so well match the evolutionary picture predicted by the theory?
Really, step away from the creationist propaganda and try reading some actual *science* for a change... You have a vast amount of misinformation in your head that can be cured by actual facts, if you just open your eyes and go looking. Little do you know how little you know.
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49
Called the information desk again about the UN
by skyman ini have called many times in the past.
this time i was calling for a sister that wanted to know the truth from the society.
the brother from the information desk said they did join but realy they were tricked.
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hawkaw
Tricked!!!!!
ROTFMAO.
Yeah right.
The problem with this argument is that each and every year after the Watchtower was approved by the DPI committee in 1992, the DPI (Department of Public Information) prescribed that the WTS along with all other "associated NGOs" must renew its associated status, fill out the prescribed paper work, sign the prescribed paperwork (done by Ciro as well as either Lloyd Barry or Don Adams) and make a note of what activities it did to dissiminate UN activities to the public. They were also provided with DPI directories that showed all the NGOs with consultative and associated status with the UN. In the preambles, hese driectories clearly explained what was expected of these organizations.
Another problem is the UN itself makes the non governmental organization (NGO) voluntarily apply to it and as part of the application process, the NGO must upfront be willing to support the UN Charter.
Enough said
hawk
http://www.geocities.com/osarsif/un.htm
http://www.un.org/dpi/ngosection/watchtower.pdf
p.s. - The WTS never joined the UN as only countries can join - even the WTS staff can't seem to get this straight. What they did was, as an existing non governmental organization, they voluntarily applied for an "associated status" with the UN's Department of Information. It was considered a partnership.
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22
Sep 15th WT: At it again: WT says elements in the UN will attack religion
by truthseeker init seems that the watchtower has not learnt any lessons regarding their involvement in the united nations.
perhaps they think the outcry that a few witnesses made a couple of years ago has died down.
they have been silent about the united nations for about 3 years until they published an article in the sep 15th watchtower, "walk by faith, not by sight!
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hawkaw
Turn the computer on and what do I read but this little gem ....
You think they would be smart and keep the UN out of their documents so people would forget but nooooooo .....
I'm going to have fun with this one.
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43
In 70 words or less - Tell one person the 'truth about the Truth'...
by AK - Jeff inwithout use of profanity, vulgarity, or sarcasm.
tell this hypothetical person who is still a dyed in the wool witness clearly why he/she should leave.
do it in such a way that he/she will not bolt for the door screaming "get behind me satan" - in other words, use tact to anti-witness.. jeff
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hawkaw
Why they should leave?
The leadership practices open bigotry; knowlingly misrepresents science to encourage members to take their lives and thus, conspires in homicide; knowingly protects child molestors and silences the children; and forces members to attend jail or face death by political groups while the leadership partners with polticial organizations such as the United Nations.
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31
Wacky Email to Quotes: why do they even bother
by Quotes inyes, that's right folks.
it is time for another peek inside the quotes mail bag.
i responded thusly:dear [name removed for privacy],.
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hawkaw
Where do these guys come from.
New interest in copyright law .... screaming when you bend over - LOL!!!!!
What a great morning laugh.
hawk